Sorry, Jack, I missed this question earlier. The light yellow-gold legs on the queen make me think she is Italian/Cordovan. Probably a mix, given her dark offspring, although even they have yellow fuzz on the thorax. I would guess the drone was Italian, contributing to their darker abdomens. Here is a very nice chart that @Valli posted last year, which gives you an idea of colouring:
Beautiful photos, as ever. You must be really good at your day job! 